I found this out first-hand when loading up my copy of Pokémon Gold—whether I’d caught ’em all back in 2001, I’ll never be able to confirm, as the battery died at some point in the last quarter-century(!). Booting it up through the GB Operator was like switching it on for the first time. However, if it had still contained my surely completed Pokédex, I’d have been able to copy that data to my laptop’s hard drive, replace the battery in the cartridge (a fiddly process, but doable), and then…

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